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Cited article:

Preserving Population Allele Frequencies in Ex Situ Conservation Programs

MARÍA SAURA, ANDRÉS PÉREZ-FIGUEROA, JESÚS FERNÁNDEZ, MIGUEL A. TORO, and ARMANDO CABALLERO
Conservation Biology 22 (5) 1277 (2008)
https://doi.org/10.1111/j.1523-1739.2008.00992.x